Syllabus Edition

First teaching 2024

First exams 2026

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Productivity During Succession (HL) (DP IB Environmental Systems & Societies (ESS))

Revision Note

Alistair Marjot

Expertise

Biology & Environmental Systems and Societies

Productivity During Succession

  • Productivity in ecosystems is the rate at which energy is produced and stored, measured as gross productivity (GP) and net productivity (NP)

    • Gross productivity is the total amount of energy captured by producers (like plants) through photosynthesis

    • Net productivity is the energy that remains after respiration by the producers, which is available for growth and consumption by other organisms

Early stages of succession (pioneer community)

  • In the early stages of succession, gross productivity is low due to:

    • Harsh environmental conditions (e.g. bare rock or poor soil)

    • Low nutrient availability (limits the growth of producers)

    • Low density of producers:

      • Pioneer species like mosses and lichens are the first to colonise these areas

      • There are fewer plants or algae to carry out photosynthesis

      • The conditions are not yet favourable for large-scale plant growth

  • Net productivity is relatively high and increases quickly at this stage because:

    • Most energy produced through photosynthesis goes toward growth and biomass accumulation

    • There is less cellular respiration taking place, as the ecosystem has fewer organisms, especially consumers

Mid-stages of succession

  • As succession progresses, conditions improve:

    • Soil formation occurs, allowing larger plants like shrubs and trees to establish

    • More nutrients become available, and the number of producers increases

  • Gross productivity rises:

    • More plants are present to carry out photosynthesis

    • Plant diversity increases, which increases the efficiency of energy capture

  • Net productivity still increases but the rate of increase slows and will eventually reach a peak

  • The rate of biomass accumulation slows because:

    • Respiration increases as more organisms, including herbivores and decomposers, become part of the system

    • There is more competition for resources like light, water, and nutrients, slowing the growth of new biomass

Late stages of succession (climax community)

  • In the final stages of succession:

    • The ecosystem reaches a stable state with a mature community of species e.g. large trees in a mature forest or diverse plant species in grasslands

    • Gross productivity may be high, as the large number of producers continue to photosynthesise

    • However, net productivity decreases and approaches zero because:

      • Most of the energy produced through photosynthesis is used up by cellular respiration of both producers and consumers

      • Biomass no longer accumulates significantly; instead, it cycles between organisms

    Graph showing changes in biodiversity, biomass, respiration, and NPP over 200 years, from pioneer to climax community. Biodiversity, biomass, and respiration plateau, while NPP declines.
    Three text boxes are connected by green arrows. They explain the stages of succession: early (low GPP, high NPP, increasing biomass), middle (increased GPP and biomass, peak NPP), and late (stable biomass with no net productivity).
    Changes occurring in a community as it develops from a pioneer community into a climax community through the process of succession

Comparison of Pioneer and Climax Communities

Pioneer Communities

Climax Communities

Stage in succession

Early stages

Later Stages

GPP

Low

High

NPP as a % of GPP

High

Low

Species richness and diversity

Low

High

Niches

Fewer, wider

Many, narrow

Size of organisms

Small

Large

Species composition

Fewer species, adapted to harsh conditions

More species, adapted to stable conditions

Total biomass (amount of organic matter)

Low

 High

Soil depth

Shallow

Deep

Soil quality

Poor (little nutrients and organic material)

High (nutrient-rich and full of organic matter)

Growth rate

Rapid

Slower

Energy flow

Simple and linear

Complex and cyclic

Nutrient cycling

Less efficient, open system (external inputs)

More efficient, closed system (nutrients are recycled)

Dominant organisms

Lichens, mosses, algae, bacteria, and fungi

Woody plants, trees, and shrubs

Stability

Unstable, prone to disturbance and colonisation

Stable, resistant to disturbance and colonisation

Examples

Pioneer species like lichens and mosses on rocks

Ancient oak forests

Examiner Tip

Don’t confuse NP with biomass; NP is the rate of energy available for growth, while biomass is the stored energy.

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Alistair Marjot

Author: Alistair Marjot

Expertise: Biology & Environmental Systems and Societies

Alistair graduated from Oxford University with a degree in Biological Sciences. He has taught GCSE/IGCSE Biology, as well as Biology and Environmental Systems & Societies for the International Baccalaureate Diploma Programme. While teaching in Oxford, Alistair completed his MA Education as Head of Department for Environmental Systems & Societies. Alistair has continued to pursue his interests in ecology and environmental science, recently gaining an MSc in Wildlife Biology & Conservation with Edinburgh Napier University.