Syllabus Edition
First teaching 2024
First exams 2026
Productivity During Succession (HL) (DP IB Environmental Systems & Societies (ESS))
Revision Note
Written by: Alistair Marjot
Reviewed by: Bridgette Barrett
Productivity During Succession
Productivity in ecosystems is the rate at which energy is produced and stored, measured as gross productivity (GP) and net productivity (NP)
Gross productivity is the total amount of energy captured by producers (like plants) through photosynthesis
Net productivity is the energy that remains after respiration by the producers, which is available for growth and consumption by other organisms
Early stages of succession (pioneer community)
In the early stages of succession, gross productivity is low due to:
Harsh environmental conditions (e.g. bare rock or poor soil)
Low nutrient availability (limits the growth of producers)
Low density of producers:
Pioneer species like mosses and lichens are the first to colonise these areas
There are fewer plants or algae to carry out photosynthesis
The conditions are not yet favourable for large-scale plant growth
Net productivity is relatively high and increases quickly at this stage because:
Most energy produced through photosynthesis goes toward growth and biomass accumulation
There is less cellular respiration taking place, as the ecosystem has fewer organisms, especially consumers
Mid-stages of succession
As succession progresses, conditions improve:
Soil formation occurs, allowing larger plants like shrubs and trees to establish
More nutrients become available, and the number of producers increases
Gross productivity rises:
More plants are present to carry out photosynthesis
Plant diversity increases, which increases the efficiency of energy capture
Net productivity still increases but the rate of increase slows and will eventually reach a peak
The rate of biomass accumulation slows because:
Respiration increases as more organisms, including herbivores and decomposers, become part of the system
There is more competition for resources like light, water, and nutrients, slowing the growth of new biomass
Late stages of succession (climax community)
In the final stages of succession:
The ecosystem reaches a stable state with a mature community of species e.g. large trees in a mature forest or diverse plant species in grasslands
Gross productivity may be high, as the large number of producers continue to photosynthesise
However, net productivity decreases and approaches zero because:
Most of the energy produced through photosynthesis is used up by cellular respiration of both producers and consumers
Biomass no longer accumulates significantly; instead, it cycles between organisms
Comparison of Pioneer and Climax Communities
Pioneer Communities | Climax Communities | |
---|---|---|
Stage in succession | Early stages | Later Stages |
GPP | Low | High |
NPP as a % of GPP | High | Low |
Species richness and diversity | Low | High |
Niches | Fewer, wider | Many, narrow |
Size of organisms | Small | Large |
Species composition | Fewer species, adapted to harsh conditions | More species, adapted to stable conditions |
Total biomass (amount of organic matter) | Low | High |
Soil depth | Shallow | Deep |
Soil quality | Poor (little nutrients and organic material) | High (nutrient-rich and full of organic matter) |
Growth rate | Rapid | Slower |
Energy flow | Simple and linear | Complex and cyclic |
Nutrient cycling | Less efficient, open system (external inputs) | More efficient, closed system (nutrients are recycled) |
Dominant organisms | Lichens, mosses, algae, bacteria, and fungi | Woody plants, trees, and shrubs |
Stability | Unstable, prone to disturbance and colonisation | Stable, resistant to disturbance and colonisation |
Examples | Pioneer species like lichens and mosses on rocks | Ancient oak forests |
Examiner Tips and Tricks
Don’t confuse NP with biomass; NP is the rate of energy available for growth, while biomass is the stored energy.
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