Efficiency of Energy Transfers
- A very large proportion of the Sun's energy is not made available to photosynthetic plants, because
- Light falls away from plants
- Light passes through leaves or is reflected away
- Light is a mixture of wavelengths, and only certain wavelengths stimulate photosynthesis
- During photosynthesis, primary producers such as plants and algae convert light energy to chemical energy in biological molecules
- The storing of this chemical energy as plant biomass makes a certain amount of energy available to the next trophic level, the primary consumers
- Only a small percentage of plant biomass becomes biomass in the primary consumer because:
- Not all the plant's biomass is eaten by the primary consumer
- Not all the consumer's biomass intake is digested
- Think about the energy content of cow dung, which can be dried and used as a heating/cooking fuel, as it contains a lot of undigested biomass e.g. cellulose
- The primary consumer converts a lot of chemical energy to movement and heat, and only a small amount to new biomass in its own body
- The efficiency of biomass transfer from one trophic level to the next is low, typically around 10%
Calculating efficiency of biomass transfers between trophic levels
- Given the appropriate data, it is possible to calculate the efficiency of biomass transfer from one trophic level to the next, as a percentage
Efficiency of transfer = (biomass transferred biomass intake) 100
- Where:
- Biomass transferred = biomass that has passed to the higher trophic level
- Biomass intake = biomass of the lower trophic level that has been consumed
Worked example
A blackberry bush with a mass of 35 kg is fed upon by aphids with a collective mass of 4.1 kg. Calculate the percentage efficiency of energy transfer in this step of the food chain.
Step 1: Ensure both units are the same
In this case, both are expressed in kg so the units do not need to be converted
Step 2: Substitute the values into the formula
Efficiency of transfer = (biomass transferred biomass intake) 100
(4.1 35) 100 % = 11.7 %
Calculating the efficiency of energy transfer between trophic levels
- A common way of working out the efficiency of energy transfer is calculating the net productivity of one trophic level as a percentage of the net productivity of the preceding trophic level
Net productivity of producers
- The net productivity of producers (also known as net primary productivity or NPP) can be calculated using the following equation:
NPP = GPP - R
- Where:
- GPP = gross primary productivity
- R = respiratory losses
Net productivity of consumers
- The net productivity of consumers can be calculated using the following equation:
N = I - (F + R)
- Where:
- I = the chemical energy store in ingested food
- F = the chemical energy lost to the environment in faeces and urine
- R = the respiratory losses to the environment
Worked example
A wheat farmer decides to use biological control against insect pests that are eating her wheat crop. The farmer introduces a species of toad. By eating the insect pests, the toads ingest 10,000 kJ m-2 yr-1 of energy but lose 2,000 kJ m-2 yr-1 of this energy in faeces and urine. They lose a further 7,000 kJ m-2 yr-1 using energy for respiration. Calculate the percentage efficiency of energy transfer from the insects to the toads.
Step 1: Calculate the net productivity of the toads
N = I - (F + R)
N = 10,000 - (2,000 + 7,000)
N = 10,000 - 9,000
N = 1,000 kJ m-2 yr-1
Step 2: Write out the equation for % efficiency and substitute in the known values
% Efficiency = (1,000 ÷ 10,000) × 100
Step 3: Calculate the efficiency
% Efficiency = (0.1) × 100
% Efficiency = 10%
How human activities can manipulate the transfer of biomass through ecosystems
- Human activity can adjust the efficiency of transfer of biomass between trophic levels, usually to maximise it in the context of maximising agricultural productivity
- For producers, arable farmers can adopt these methods
- Providing artificial light in greenhouses on overcast days
- Optimising planting distances between crops
- Irrigation to maximise growth in dry weather
- Use of fertilisers
- Selective breeding for fast growth
- Use of fungicides/pesticides
- Fencing to exclude grazers
- Ploughing and herbicides to kill weeds
- Plant crops that store energy in edible form e.g. seeds, fruit, tubers
- Livestock farmers can adopt these methods for primary consumers (grazers)
- Use of good quality feeds / food supplements
- Use antibiotics and vaccines to reduce disease
- Control predation with fencing or with indoor animal husbandry
- Reduce competition for grazing e.g. rabbits, deer
- Indoor husbandry to reduce energy loss from movement or from getting cold outside
Human activity can increase the efficiency of biomass transfer in an arable farm setting
Examiner Tip
Exam questions can refer to biomass and energy interchangeably. Remember, the biomass of an organism is effectively a measure of how much chemical energy is stored within it!