Phloem Sieve Tube Elements & Companion Cells: Structure & Function
- The function of phloem tissue in a plant is to:
- Transport organic compounds (assimilates), particularly sucrose, from the source (eg. leaf) to the sink (eg. roots). The transport of these compounds can occur up and down the plant
- The organic compounds are dissolved in water to form sap
- Phloem is a complex tissue made up of various cell types; its bulk is made up of sieve tube elements which are the main conducting cells and companion cells
- Other cell types of phloem tissue also include parenchyma for storage and strengthening fibres
- Mature phloem tissue contains living cells, unlike xylem tissue
Structure of phloem tissue. (a) Microscope slide image and drawing of a sieve tube element and companion cell in transverse section (TS), (b) photomicrograph image and drawing of a sieve tube element and companion cell in longitudinal section (LS).
Structure of phloem tissue. (c) Transmission electron micrograph image and drawing of a sieve tube element and companion cell in transverse section (TS)
Sieve tube elements
- Sieve tube elements line up end to end to form a continuous tube
Phloem sieve tube elements structure & function table
Companion cells
- Each sieve tube element has a companion cell associated with it as companion cells control the metabolism of their associated sieve tube member
- They also play a role in loading and unloading of sugars into the phloem
Phloem companion cells structure & function table
Comparison of xylem & phloem tissue table
Examiner Tip
Understand the difference between sieve tube elements and companion cells, and how they are different to xylem tissue. Remember that mature xylem tissue is dead, so there is no evidence of organelles, and they have lignified cell walls, whereas sieve tube elements have no lignin, do have sieve plates, and their companion cells contain nuclei and dense cytoplasm.